The fight rages between Doma, the 2nd upper moon and the gunners Kanao and Inosuke. Will the two young warriors, trembling with anger, manage to avenge all those whom Doma devoured? Not far from there in the Infinite Fortress, Mitsuri Kanroji, the pillar of Love, and Obanai Iguro, the pillar of the Serpent, face the new 4th upper moon, while Muichiro Tokito, the pillar of the mist, finds himself facing the 1st upper moon, which makes him a strange revelation about his family...